A penguin has died at the Calgary Zoo.

Guillermo, a three-year-old Humboldt penguin, came to Calgary in 2011 from a zoo in New York.

Zoo keepers say he was lethargic and didn't want to eat over the weekend.

Story continues below advertisement

He died Monday night.

The penguins are one of the most popular attractions at the zoo, which recently reopened after suffering heavy damage during the June floods.